Borrego Locoweeds
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Locoweeds Of Borrego Valley & Vicinity (Anza Borrego State Park)
(Images Of Several Other Species Outside Of Borrego Valley Are Also Included)
© W.P. Armstrong 16 March 2010

Locoweeds belong to the large and diverse genus of flowering plants Astragalus (Fabaceae), with approximately 2,000 different species in the northern hemisphere, 380 species in North America, and 94 species in California. Some estmates are over 3,000 species! Some are rare endemics that only occur in one isolated mountain range or canyon. They are sometimes called milk vetches from the notion that milk secretion in goats was increased when they fed on the common Old World forage species (Astragalus cicer). Like many legumes, the leaves are typically divided into a dozen or more leaflets, and the flowers resemble small pea blossoms. Some species produce inflated seed pods that make a distinct popping sound if you step on them. Locoweeds are also called rattleweed because of the bladder-like, inflated pods, particularly when a gust of wind rattles the seeds inside. Astragalus is derived from a Greek word meaning anklebone, the plural of which means dice. Perhaps the dice connotation refers to the rattling of seeds inside papery pods, like the sound of dice in a thrower. In anatomy, the astragalus or talus is one of seven bones in the ankle joint. Anklebones were apparently used for dice by ancient Greeks, and to this day, veteran crapshooters in Las Vegas refer to dice as "bones." Since adult astragalus bones are a little too large for dice, some of the smaller, cuboidal or cuneiform anklebones were probably used.


Largest Genera Of Flowering Plants With 800 or More Species*

FAMILY
GENUS
# OF SPECIES
ARACEAE
Anthurium
900
ASTERACEAE
Senecio
Vernonia
1250
1000
BEGONIACEAE
Begonia
919
CYPERACEAE
Carex
2000
ERICACEAE
Erica
Rhododendron
860
850
EUPHORBIACEAE
Euphorbia**
Croton
2400
1300
FABACEAE
Astragalus***
Acacia
2000 (3000!)
1000
LAMIACEAE
Salvia
800
MELASTOMATACEAE
Miconia
1000
MORACEAE
Ficus
800+
ORCHIDACEAE
Bulbophyllum
Dendrobium
Epidendrum
Pleurothallis
1000
900
800
1120
OXALIDACEAE
Oxalis
800
PHYLLANTHACEAE
Phyllanthus
1270
PIPERACEAE
Peperomia
1000
ROSACEAE
Rosa
2050
RUBIACEAE
Psychotria
1500
SOLANACEAE
Solanum
1400

* Judd, W.S., Campbell, C.S., Kellogg, E.A., Stevens, P.F., and M. J. Donoghue. 2008. Plant Systematics:
A Phylogenetic Approach
(Third Edition). Sinauer Associates, Inc., Sunderland, Massachusetts.

** Some authors subdivide Euphorbia into several genera incl. the prostrate sandmats (Chamaesyce).
The On-line Plant List By Kew & Missouri Botanical Garden gives 2031 species for Euphorbia.

*** The On-line Plant List By Kew & Missouri Botanical Garden gives 2,481 accepted species
for the genus Astragalus, 2029 species for Carex, and 149 species for Rosa.

Working List Of All Known Plant Species. 2011. Kew & Missouri Botanical Garden.
Publishished on the Internet: www.theplantlist.org/

In other words, if all the 250,000 species of flowering plants on earth were lined up at random, every 4th species would be a sunflower (Asteraceae), a legume (Fabaceae) or an orchid (Orchidaceae)! If you happen to pick a legume, the odds that it will be a locoweed (Astragalus) is about one out of nine.

Barneby, R.C. 1964. "Atlas of North American Astragalus." Volumes I & II.
Memoirs of the New York Botanical Garden 13: 1-1188.


Astragalus lentiginosus var. borreganus, an endemic locoweed in the Borrego Valley area.

Desert dwarf locoweed Astragalus didymocarpus var. dispermus, a common prostrate annual in sandy areas.

Small seed pods of Astragalus didymocarpus var. didymocarpus.

  Size Of Sewing Needle Used In Wayne's Word Images  

Astragalus aridus, an uncommon annual locoweed with silvery appressed hairs.

Sand verbena (Abronia villosa var. villosa) & seed pods of Astragalus aridus (right).

Astragalus palmeri, a perennial locoweed native to the Anza-Borrego Desert region. Unlike the erect habit of Astragalus lentiginosus var. borreganus, this species typically has a low, spreading growth form.

Astragalus crotalariae, a perennial locoweed in the badlands east of Borrego Springs. The peculiar malodorous scent of dried specimens lasts for years in herbarium cabinets. It is one of the species of locoweeds known to concentrate toxic levels of selenium.

Astragalus nuttallianus var. cedrosensis: A small annual locoweed.
(Photo of herbarium specimen just taken out of plant press.)

Astragalus insularis var. harwoodii (Harwood's Rattleweed).


Images Of Several Other Species Outside Of Borrego Valley

Astragalus magdalenae var. piersonii (dune locoweed): A rare endemic native to the Algodones Sand Dunes.

Astragalus douglasii ssp. parishii on the crest of the Laguna Mts. overlooking Anza-Borrego Desert. This perennial locoweed has inflated, bladderlike pods that rattle in the wind.

Astragalus leucolobus: A rare perennial with fury pods native to the San Gabriel Mountains.

Scarlet locoweed (Astragalus coccineus) from Mountain Springs Grade.

Astragalus funereus: A rare endemic to the Funeral Mountains of eastern Death Valley.

A Locoweed & A Dead Steer
Mind Altering Plant Alkaloids
Model Of Locoweed Poison Molecule
Endemic Dune Locoweed & Sand Food
  Locoweed Natural Gum & Medicinal Herb  


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